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Finally, lysosomal and liposomal colocalization was imaged with a high-content screening system in confocal mode, and Pearson correlation coefficients were calculated by using GraphPad Prism 8.3.0 software 67

By inhibiting these enzymes, TIMP-4 helps to regulate the ECM turnover and tissue remodelling in the heart and the brain [134,135,136] (Table 1)

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