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Data availability The cryo-EM maps have been deposited in the Electron Microscopy Data Bank (EMDB) under the accession codes EMD-71735 (OCTN2 inward-facing), EMD-71540 (OCTN2 occluded) and EMD-71597 (OCTN2 ipratropium)

The true body weight was calculated by subtracting the tumor weight from the total body weight

Meanwhile, for colonic goblet cells, which are another type of secretory cell, their impaired differentiation and function have been associated with UC (9799)

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Nuclear staining identifies two populations of human sperm with different DNA fragmentation extent and relationship with semen parameters

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