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Second, normalization to the mitochondrial matrix enzyme citrate synthase (CS), an established marker for mitochondrial content, enabled us to distinguish whether the increase in OXPHOS proteins was due to an overall rise in mitochondrial content or represented a selective upregulation of the OXPHOS complexes on the mitochondrial membrane relative to other mitochondrial proteins

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doi: 10.1186/s12967-016-0771-6 93 VenterMTomasCPienaarISStrassheimVErasmusENgWFet al